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Mrs. Jeffries is the ex-wife of Arthur Jeffries aka Professor Proton and mother of their son.

It is revealed in "The Proton Resurgence" that the the puppeteer who did Gino the Neutrino also did Mrs. Jeffries (made a move on her) and this caused Arthur to hate the puppet Gino.

In "The Proton Transmogrification", Sheldon asked Arthur's spirit why he is in Apartment 4A and Arthur replies that he doesn't know and then says maybe come back to haunt his ex-wife.